A spokesman for the U.S. State Department said no agreement on such action has been made with Russia.

Defense Minister Sergei Shoigu said that Moscow had proposed to Washington that the coalition call on all factions eager to abide by a cease-fire in Syria to leave the areas where the al-Qaida branch is active by May 25. Then Russia and the U.S.-led coalition could conduct joint strikes against the Nusra Front and any other groups refusing to honor the truce, Shoigu suggested.

State Department spokesman John Kirby said "there is no agreement to conduct joint airstrikes with the Russians in Syria" but added Washington and Moscow are discussing "proposals for a sustainable mechanism to better monitor and enforce the cessation of hostilities."

Earlier, Pentagon spokesman Navy Capt. Jeff Davis would not comment directly on the proposal, which he said has not been formally presented to the United States. But he said the U.S. military is not cooperating or collaborating on operations with Russia in Syria except to maintain airspace safety.

"Russian operations are supporting the Assad regime and our focus is solely on degrading and defeating ISIL," he said, using an alternative acronym for the Islamic State.

Shoigu warned that Moscow reserves the right to unilaterally strike militants refusing to respect the cease-fire, as well as weapons and militants crossing into Syria from Turkey, starting from May 25.

Russia long has accused Turkey of serving as a conduit for weapons and supplies flowing to the Islamic State and the Nusra Front — claims which Ankara has denied.

The U.S.-Russia-brokered cease-fire, which went into effect in late February, has helped reduce violence in Syria, but it has been steadily eroding. The Nusra Front and its much more powerful rival, the Islamic State group, have been excluded from the cease-fire, and Russia long has pushed for excluding other militant groups in Syria that have cooperated with the al-Qaida affiliate.

Shoigu said the Russian proposal should help secure the cease-fire, adding that Moscow has coordinated it with Damascus. He claimed that the Russian military has started discussing the idea with the U.S. representatives.

Meanwhile in Syria, rebels and pro-government forces fought on several fronts on Friday after the Syrian army, backed by the Lebanese Hezbollah militant group, managed to seize valuable territory around the capital, Damascus.

Activists reported continued fighting in the Ghouta suburbs of Damascus with warplanes — belonging to either Russia or the Syrian military — intensifying airstrikes on the besieged, rebel-held area of Daraya.

Airstrikes on a rebel-held village in the northwest Idlib province killed at least eight civilians, according to the Local Coordination Committees group. The Britain-based Syrian Observatory for Human Rights, another group that tracks the Syrian war, said nine had died, including two children and three women.

The Observatory also raised its death toll from airstrikes Thursday on rebel-held villages in the central Homs province to 22 civilians.

Since the Russian military campaign began last September in an effort to shore up Syrian President Bashar Assad's forces in their battle against Islamic militants, Moscow has staunchly denied that its warplanes have hit any civilian areas in Syria.

Also Friday, ultraconservative rebel factions and government forces clashed in the central Hama province around the predominantly Alawite town of Zaara, which al-Qaida-linked militants seized from government control last week. The Observatory said both sides suffered casualties in the fighting.

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Associated Press Writers Deb Riechmann in Washington, Matthew Lee in Brussels and Philip Issa in Beirut contributed to this report.

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